Initial inoculum

The size of the parasite population at the beginning of the epidemic. Other things being equal, a high initial inoculum leads to a more rapid development of the epidemic, while a low initial inoculum leads to a slower or later development of the epidemic. Injury

The injury from parasitism is the actual amount of damage done to an individual host, or the average amount done to a host population, by the parasite. The frequency of parasitism is the proportion of host individuals that are parasitised. In a wild plant pathosystem, the injury from parasitism is inversely proportional to the frequency of parasitism. That is, the higher the frequency, the lower the injury, and, conversely, the higher the injury, the lower the frequency. In this way, the total damage from parasitism never exceeds a tolerable level that does not impair the host's ability to compete ecologically and evolutionarily. Vertical resistance, with its system of locking, reduces the frequency of parasitism. horizontal resistance, as a second line of defence, reduces the injury from parasitism. Continuous plant pathosystems, that have horizontal resistance only, usually have a high frequency of parasitism, and a low injury from parasitism.

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