The inverse relationship between vertical resistance and horizontal resistance

There is an inverse relationship between vertical resistance and horizontal resistance that is essentially the inverse relationship between frequency and injury (see 10.14).

If there is a vertical subsystem with a complex system of locking, the frequency of matching allo-infection is low, and the frequency of parasitism is also low. There is then very little selection pressure for horizontal resistance, which will also be at a low level, and this will result in a relatively high rate of injury.

If, however, there is a vertical subsystem with a simple system of locking (or no vertical subsystem at all), the frequency of matching allo-infection will be relatively high, and the frequency of parasitism will also be high. There will then be a strong selection pressure for horizontal resistance, which will also be high, resulting in a low rate of injury. The vertical subsystem thus determines the frequency of parasitism which, in its turn, determines the selection pressure for horizontal resistance.

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