The Families And Genera Of Vascular Plants

Edited by K. Kubitzki

Springer

Berlin Heidelberg New York HongKong London Milan Paris Tokyo

Volumes published in this series:

Volume I Pteridophytes and Gymnosperms

Edited by K.U. Kramer and P.S. Green (1990) Date of publication: 28.9.1990

Volume II Flowering Plants. Dicotyledons. Magnoliid, Hamamelid and Caryophyllid Families Edited by K. Kubitzki, J.G. Rohwer, and V. Bittrich (1993) Date of publication: 28.7.1993

Volume III Flowering Plants. Monocotyledons: Lilianae (except Orchidaceae) Edited by K. Kubitzki (1998) Date of publication: 27.8.1998

Volume IV Flowering Plants. Monocotyledons: Alismatanae and Commelinanae (except Gramineae)

Edited by K. Kubitzki (1998) Date of publication: 27.8.1998

Volume V Flowering Plants. Dicotyledons: Malvales, Capparales and Non-betalain Caryophyllales

Edited by K. Kubitzki and C. Bayer (2003) Date of publication: 12.9.2002

Volume VI Flowering Plants. Dicotyledons: Celastrales, Oxalidales, Rosales, Cornales, Ericales Edited by K. Kubitzki (2004)

The Families and Genera of Vascular Plants

Edited by K. Kubitzki

Flowering Plants • Dicotyledons

Celastrales, Oxalidales, Rosales, Cornales, Ericales

Volume Editor: K. Kubitzki

With 137 Figures

Springer

Professor Dr. Klaus Kubitzki Institut für Allgemeine Botanik Ohnhorststraße 18 22609 Hamburg Germarry

ISBN 3-540-06512-1 Springer-Verlag Berlin Heidelberg New York

Library of Congress Catalo ging-in-Publication Data

Flowering plants, Dicotyledons: Celastrales, Oxalidales, Resales, Cornales, Ericales / volume editor, K. Kubitzki. p. cm. - (Families and genera of vascular plants; 6) Includes bibliographical references (p.). ISBN 3-540-06512-1 (alk. paper)

1. Dicotyledons - Classification. L Kubitzki, Klaus, 1933- II. Series. QK495.A12F56 2004

583'.01'2 - dc22 2003058977

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Preface

During recent years the findings of molecular systematics for the first time have provided a reliable lead for a higher order classification of angiosperms, as reflected in the classification of the Angiosperm Phylogeny Group. From Volume III of this series onward, these new concepts have been used as a guideline for treating angiosperm families in a meaningful context within individual volumes of this series. Among the five orders treated in this volume, Celastrales and Oxalidales represent novel concepts of which at least the latter makes sense also in morphological terms. The concept of Ericales followed here has largely been expanded beyond its conventional limits to include parts of former Ebenales, Lecythidales, Primulales and other orders. Also, the circumscription of many families has been revised on the basis of recent molecular studies. Among the recircumscribed families, Ericacae, Celastraceae, Cunoniaceae and Elaeocarpaceae appear now in revised and expanded circumscriptions, and the former Primulales families have been profoundly reorganised. I am most grateful to my authors who have incorporated into their treatments the most recent results of molecular and cladistic studies. At the same time I, as the editor, also feel the dilemma of prejudicing those authors who responded at an early stage of the preparation of these volumes and whose contributions had to wait for publication until late authors were able to comply. Lamentably, R.C. Carolin's treatments of Stackhousiaceae and Tremandraceae, submitted in the early 1990s, have been superseded by later developments, and I am most grateful to Roger Carolin for his understanding and generous permission to use his treatments in a changed family context.

I would like to direct the attention of the users of this series to the remarkable treatment of Ericaceae in this volume. Its author, P.F. Stevens, is introducing here an interesting kind of "hierarchical" description, which emphasises apomorphic characters and leads to great clearness and conciseness, the constantly aspired aims of this series.

It would hardly have been possible to complete this volume without the help so generously offered by many colleagues. Most substantial support has been provided by C. Bayer and P.F. Stevens who reviewed many contributions and suggested most helpful improvements. A.S. George is thanked for building up a team that contributed Styphelioideae and thus made possible completion of Ericaceae in the new circumscription; I am also grateful for his kind assistance in various other matters. Deep thanks for reading and commenting on particular contributions are due to R. Archer (Celastraceae), I.K. Ferguson (Cornaceae), J.W. Kadereit (Introduction), S.R. Manchester (Cornaceae), M. Matthews (Cunoniaceae), B. Stahl (Symplocaceae), S. Renner (Sarraceniaceae), R. Rutishauser (Cunoniaceae) and J. Thompson (tremandraceous Elaeocarpaceae). D. Albach and L. Hufford are thanked for their comments on problems concerning Hydrostachyaceae, and M. Matthews and P.K. Endress for showing me papers prior to publication.

My thanks go also to the copyright holders of the illustrations who so generously authorised the use of the material included in this volume.

Finally, it is a pleasure to acknowledge the agreeable collaboration with the staff of Springer-Verlag and ProEdit, Heidelberg, who kindly responded to all requests I had in connection with the completion of this volume.

Hamburg, October 2003

K. Kubitzki

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